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Front cover: Models for cohesion establishment. Pds5 (blue) inhibits cohesin complex (red). Ctf7/Eco1 (yellow) binds to and displaces Pds5 to unlock cohesin rings. (A) One cohesin ring around both sisters. Ctf7 acts on cohesin to open ring and then Pdf5 locks cohesin again. (B) Interaction of two cohesin rings with around each sister. (C) Cohesin ring associates with each sister chromatid. In (B)and (C), Ctf7 alters cohesin to promote dimerization, then Pdf5 protects cohesin associations. For detail see the review article by V. Guacci in this issue (pp.693–708).